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Naughty doodles, split ink and inscriptions are all revealed in a new exhibition looking at the first ever printed books.
Private Lives of Print from the University of Cambridge Library displays books printed between 1450-1550. The exhibition draws on the university's collection of incunabula, which are books printed before 1501. Some are beautifully preserved, others have been scribbled on and marked:
All images courtesy of University of Cambridge